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Course Descriptions

COLLEGE OF OCEAN & FISHERY SCIENCES
OCEANOGRAPHY

Detailed course offerings (Time Schedule) are available for

To see the detailed Instructor Class Description, click on the underlined instructor name following the course description.

OCEAN 100 Explore Oceanography at UW (1)
Explores the research and experiences of the Oceanography department's faculty, graduate students, and undergraduate students. Credit/no credit only. Offered: A.

OCEAN 101 Survey of Oceanography (5) NW
Holistic view of fundamental principles of ocean science; the geography and geology of ocean basins; chemistry of sea water; physical dynamics of currents, waves, and tides; coastal processes; and the biology of diverse ecosystems such as deep sea vents, coral reefs, and estuaries. Intended for nonmajors.
Instructor Course Description: Richard M. Strickland

OCEAN 102 The Changing Oceans (5) I&S/NW
Historical case studies of research on the ancient oceans, deep-sea exploration, climate change and the oceans, and human impacts on marine life. Students consider societal factors affecting progress in marine science, changing popular attitudes toward the oceans, and key current policy implications of marine science. Intended for nonmajors. Offered: W.
Instructor Course Description: Richard M. Strickland

OCEAN 115 Astrobiology: Life in the Universe (5) NW
Introduction to the new science of astrobiology, study of the origin and evolution of life on Earth, and the search for microbial and intelligent life elsewhere in the Universe. Designed for non-science, liberal arts majors. Offered: jointly with ASTBIO/ASTR/ESS 115/BIOL 114.

OCEAN 200 Introduction to Oceanography (3) NW Armbrust
Description of the oceans. Emphasis on relations of biology, chemistry, geology, and physics in marine environments. Examination of relationships and interactions at macro-, meso-, and microscales in the ocean. Intended for science majors. Offered: Sp.
Instructor Course Description: E. Virginia Armbrust

OCEAN 210 Ocean Circulation (3) NW
The large-scale circulation of the ocean. Topics include temperature-salinity analysis; water mass identification; water, salt, and heat budgets; chemical tracer distributions; advection and diffusion. Prerequisite: either Ocean 101, Ocean 200, or OCEAN 250/BIOL 250/FISH 250; recommended: either PHYS 114 or PHYS 121. Offered: A.
Instructor Course Description: Paul D Quay

OCEAN 220 Introduction to Field Oceanography (3/5) NW
Design and conduct a field study in oceanography. Field trip required (usually during Spring break). Focus on active learning, deployment of instruments, data collection, interpretation, and presentation. Honors section incorporates additional field experimentation and study in marine biology. Writing class. Prerequisite: either OCEAN 101 or OCEAN 200; recommended: OCEAN 210. Offered: Sp.
Instructor Course Description: Gabrielle L. Rocap

OCEAN 230 Rivers and Beaches (3/5) NW Montgomery, Nittrouer
Introduction to Earth surface environments, the processes that shape them, how humans affect them and are affected by them. Weekend field trips examine mountains, rivers, deltas/estuaries, beaches, and environments beyond. Focus on linkages between these environments to illustrate coupling between landscapes and seascapes. Offered: jointly with ESS 230.

OCEAN 240 Contemporary Issues in Oceanography (1-5, max. 9) NW
Selected topics of contemporary interest in oceanography such as hydrothermal vents, planetary volcanism, biogeochemical cycling, the ecology of Puget Sound, and the ocean's role in climate.
Instructor Course Description: Neil S. Banas

OCEAN 250 Marine Biology (3/5) I&S, NW
Lecture-laboratory course in Marine Biology focusing on physical, biological, and social aspects of the marine environment. Topics include oceanography, ecology, physiology, behavior, conservation, fisheries, exploration, and activism. Evening marine biology movies and weekend field trip. Honors section research project. Offered: jointly with BIOL/FISH 250.
Instructor Course Description: M. Claire Horner-Devine

OCEAN 260 The Puget Sound Ecosystem (3/5) I&S/NW
Examines historical human impacts on the land/water ecosystem of the Puget Sound, roles of regional governance and citizen action, and prospects for ecological restoration. Computer labs and field trips for additional credit. Offered: jointly with ENVIR 260.
Instructor Course Description: Richard M. Strickland

OCEAN 300 Study Abroad Marine Sciences (1-12, max. 12) Robigou
For participants of Marine Language Exchange Scholarship Program. Specific content varies and must be individually evaluated. Credit does not apply to major requirements without approval. Offered: AWSpS.

OCEAN 310 The Puget Sound Nearshore: Processes and Problems (3) I&S/NW
An introduction to the geology, ecology, and politics of Puget Sound beaches. Content focuses on beach formation and sediment dynamics, with application to nearshore ecology and shoreline management. Three filed trips to local beaches complement topics discussed in class.
Instructor Course Description: Jeffrey D Parsons

OCEAN 351 Field Investigations in Marine Biology (5)
Evaluates the relationships between man and marine systems in a field-oriented class. Case studies directly investigate marine biology. Studies include human activities and our effects on marine species and communities. Multiple field trips, lectures, and labs. Prerequisite: FISH 350/OCEAN 350. Offered: jointly with FISH 351; Sp.
Instructor Course Description: Richard G Keil

OCEAN 355 From Big Bang to the Blue Planet (3) NW Sachs
Explores the origin and evolution of the Earth, ocean, atmosphere, and life, with an emphasis on climate as the integrator of changes in the biosphere, cryosphere, hydrosphere, atmosphere, and lithosphere. Prerequisite: either CHEM 120 or CHEM 142; either PHYS 114 or PHYS 121; either BIOL 161, or BIOL 180.
Instructor Course Description: Julian P. Sachs

OCEAN 360 Marine Field Research (10) NW,QSR Scott Veirs, Val Veirs
Intensive off-campus marine research experience. Includes 5 weeks designing a field experiment and a 5-week cruise aboard a sailing research vessel to implement it. Offered: jointly with OCEAN 365; ASp.

OCEAN 365 Practicing Sustainability Science (8) I&S/NW Scott Veirs, Val Veirs
Intensive off-campus experience. Studies sustainability science, marine policy, and "clean" technologies. Includes interaction with community stakeholders for 5 weeks and experimentation with sustainable technologies and practices in the marine environment during a 5-week cruise aboard an energy-efficient sailing catamaran. Offered: jointly with OCEAN 360; ASp.

OCEAN 400 Chemical Oceanography (4) NW
Physical and chemical properties of seawater and marine products; processes determining the chemical makeup of the oceans. Prerequisite: either CHEM 152 or CHEM 155; OCEAN 210. Offered: W.
Instructor Course Description: James W Murray

OCEAN 401 Special Topics in Chemical Oceanography (3) NW

OCEAN 410 Marine Geology and Geophysics (4) NW
Sedimentological and petrologic processes that determine the geologic record. Prerequisite: either ESS 101 or ESS 210. Offered: jointly with ESS 410; A.
Instructor Course Description: William S D Wilcock

OCEAN 411 Special Topics in Marine Geology and Geophysics (3) NW
.

OCEAN 420 Physical Processes in the Ocean (4) NW
Physical properties and processes of the ocean: methods of describing ocean currents, waves, tides and mixing and their effect on movement of water parcels. Prerequisite: either PHYS 116 or PHYS 123; either MATH 126, MATH 129, MATH 146, or Q SCI 381; OCEAN 210. Offered: W.

OCEAN 421 Special Topics in Physical Oceanography (3) NW

OCEAN 422 Ocean Dynamics (3) NW
Equations of motion governing flow of sea water. Conservation of mass, tracers, heat and momentum. Energy and vorticity balance. Buoyancy and rotational effects. Scale analysis. Applications to upper ocean dynamics, surface and internal waves, and wind-driven currents in the ocean. Prerequisite: MATH 126; PHYS 123; OCEAN 420.

OCEAN 423 Ocean Circulation and Climate (3) NW
Quantitative treatment of ocean basin to global scale ocean circulation systems and their interaction with climate variability. Prerequisite: PHYS 123; MATH 125; recommended: OCEAN 410.

OCEAN 430 Biological Oceanography (4) NW
Marine organisms, their quantitative distribution in time and space and their interactions with the ocean. Prerequisite: either BIOL 162 or BIOL 200; OCEAN 210; recommended BIOL 220. Offered: A.
Instructor Course Description: Gabrielle L. Rocap

OCEAN 431 Special Topics in Biological Oceanography (3) NW

OCEAN 442 Oceanography of the Puget Sound (3) NW
Explores the role of oceanography in regional issues. Field opportunities and active investigation of applied oceanographic problems. Lectures, research trip, student co-teaching, discussion. Prerequisite: either CHEM 152, CHEM 221, BIOL 162, BIOL 220, ESS 101, or ESS 210.. Offered: A.
Instructor Course Description: Richard G Keil

OCEAN 443 Design of Oceanographic Field Experiments (3) NW
Case histories, presentations, and class exercises used to teach methods of formulating a research problem and proposal writing. Methods of data analysis, presentation, error estimation, library resource and data base use; web page implementation and design. Principles of cruise planning. Prerequisite: OCEAN 400; OCEAN 410; OCEAN 420; OCEAN 430. Offered: W.

OCEAN 444 Advanced Field Oceanography (5) NW
Conduct field experiment (designed in OCEAN 443) during a week-long cruise aboard a research vessel. Analyze samples data and present results in a series of drafts and a final term paper. Results are presented at a two-day-long public research symposium and on the students' individual Web sites. Prerequisite: OCEAN 443. Offered: Sp.

OCEAN 450 Climatic Extremes (4) NW
Course examines earth history for extreme climatic conditions to predict future climate changes. Numerical climate models use PC-based computer programs to identify processes and feedbacks that control climate
Instructor Course Description: Harlan Paul Johnson Paul D Quay

OCEAN 451 Fluid Dynamics Laboratory (4) NW
Individual projects in experimental fluid dynamics with applications to practical problems. Experimental design, visualization, and measurement techniques applied to a problem selected by each student. Prerequisite: PHYS 123.

OCEAN 452 Spatial information Technologies in Ecosystem Sciences (3) NW Logsdon
Introduction to the use of GPS, GIS, and Remote Sensing in the ecosystem sciences. Integrates these technologies in an applied research setting. Two overnight weekend field trips required. Offered: jointly with FISH 453; A.
Instructor Course Description: Miles G. Logsdon

OCEAN 454 Hydrothermal Systems: An Interdisciplinary View (3) NW Lilley
Provides a general, interdisciplinary overview of seafloor hydrothermal systems including important geological, chemical and biological processes. Topics include tectonic and volcanic controls on hydrothermal systems, water/rock reactions, phase separation, temporal variability, fluxes to the deep sea, micro- and macro biology. Offered: Sp.

OCEAN 479 Research in Marine Biology (1-15, max. 15)
Individual research on topics in marine biology. Research projects supervised by an individual faculty member. Projects may include laboratory work, fieldwork, and literature surveys. Prerequisite: BIOL 250/FISH 250/OCEAN 250; Q SCI 381. Offered: jointly with BIOL 479/FISH 479; AWSpS.

OCEAN 492 Friday Harbor Apprenticeship (9/15) NW
Intensive, full-time research training experience where teams of students work on focused research problems guided by a group of faculty, postdoctoral, and graduate student mentors. Research questions vary.

OCEAN 494 Field Experiences in Marine Science (1-15, max. 30) NW faculty
For participants in oceanography field work. Specific content varies and is individually evaluated. Credit does not apply to major requirements without approval.

OCEAN 496 Study Abroad: Oceanography (1-15, max. 30) NW
For participants in UW study abroad program. Specific content varies and is individually evaluated. Credit does not apply to major requirements without approval.

OCEAN 497 Advanced Special Topics in Oceanography (1-15, max. 15)
.

OCEAN 499 Undergraduate Research (1-15, max. 24)
Research on assigned topics that may involve laboratory work, fieldwork, or literature surveys. Offered: AWSpS.
Instructor Course Description: Miles G. Logsdon Daniel Grunbaum

OCEAN 500 Current Problems in Oceanography (1)
Discussion of research topics that are currently being investigated within the school. Credit/no credit only. Prerequisite: permission of instructor. Offered: AW.

OCEAN 501 Estuarine Circulation and Mixing (3)
Observed patterns of currents, mixing, and stratification from deep fjords to shallow coastal plain estuaries. Physical understanding of basic processes, such as tides, wind stress, topographic effects on turbulence, sill hydraulics, and exchange flow. Vertical mixing and residence times important to biological and pollution studies. Prerequisite: permission of instructor.

OCEAN 506 Interdisciplinary Seminar in Oceanography (1-3, max. 12)
Lectures, discussions, and work on selected problems of an interdisciplinary nature. Prerequisite: permission of instructor.
Instructor Course Description: Eleanor E. Williams Miles G. Logsdon

OCEAN 509 Seminar (1)
Introduction to current research topics for beginning graduate students. Credit/no credit only. Offered: AWSp.

OCEAN 510 Physics of Ocean Circulation (3)
Structure of ocean basins; physical properties of seawater and the equation of state; heat, salt, fresh water budgets; tidal potential; Coriolis effect and geostrophic balance; major current systems and water masses; mixing, stirring in the ocean; simple waves; modern experimental methods in physical oceanography. Prerequisite: permission of instructor. Offered: A.

OCEAN 511 Introduction to Fluid Dynamics (4)
Eulerian equations for mass-motion; Navier-Stokes equation for viscous fluids, Cartesian tensors, stress-strain relations; Kelvin's theorem, vortex dynamics; potential flows, flows with high-low Reynolds numbers; boundary layers, introduction to singular perturbation techniques; water waves; linear instability theory. Prerequisite: AMATH 403 or permission of instructor. Offered: jointly with AMATH/ATM S 505; A.
Instructor Course Description: Dale R Durran Parker Mac Cready

OCEAN 512 Geophysical Fluid Dynamics I (4)
Dynamics of rotating stratified fluid flow in the atmosphere/ocean and laboratory analogues. Equations of state, compressibility, Boussinesq approximation. Geostrophic balance, Rossby number. Poincare, Kelvin, Rossby waves, geostrophic adjustment. Ekman layers. Continuously stratified dynamics: Inertia-gravity waves, potential vorticity, quasigeostrophy. Prerequisite: OCEAN 511 or ATM S/AMATH 505. Offered: jointly with ATM S 509; W.

OCEAN 513 Geophysical Fluid Dynamics II (3)
Theories, models of large-scale dynamics of oceans, atmospheres. Potential vorticity, Q principles; Rossby waves, ray tracing, Green's function, setup of general circulation; atmospheric "channels" versus ocean "basins"; wave-mean flow interaction, mountain drag, internal momentum flux; "Lagrangian" motion of particles, tracers; cascades, eddy flux of heat, moisture, Q. Prerequisite: OCEAN 512. Offered: Sp.

OCEAN 514 Waves (3)
Application of marine hydrodynamics principles to wave motion in oceans. Offered: W.

OCEAN 515 Ocean Circulation: Observations (3)
Modern large- and mesoscale ocean observations, interpreted in terms of contemporary circulation theories. Spectrum of temporal variability; eddies and eddy fluxes; ventilation; advection and diffusion in the abyss; transports of heat and salt; climatic scale of variability; modern methods for determining circulation. Prerequisite: OCEAN 510 or permission of instructor. Offered: Sp.

OCEAN 516 Ocean Circulation: Theories (3)
Hydrodynamic theories concerning origin and characteristics of major ocean currents. Prerequisite: OCEAN 512 or permission of instructor.

OCEAN 517 Methods and Measurements in Physical Oceanography (2)
Principal instruments and experimental methods of physical oceanography. Devices and systems that measure pressure, temperature, electrical conductivity, sea state, and velocity. Prerequisite: permission of instructor. Offered: alternate years.
Instructor Course Description: Thomas B Sanford

OCEAN 519 Seminar in Physical Oceanography (1, max. 9)
Discussion of selected problems of current interest in physical oceanography. Prerequisite: OCEAN 510 or permission of instructor.

OCEAN 520 Marine Chemistry (3)
Processes controlling the chemical composition of seawater. Chemical distributions in the ocean, marine physical chemistry, chemical equilibrium, and concepts of mass balance. Mechanisms and models used to explain distributions of stable and radioactive isotopes, gases, trace metals, and biochemicals in the world's oceans. Offered: A.

OCEAN 521 Aquatic Chemistry (3)
Application of physical chemistry and thermodynamics to processes that control chemical composition of natural waters. Equilibrium approach. Acid/base chemistry, the carbonate system, dissolution and precipitation, metal ions in solution, oxidation-reduction chemistry, silicate mineral reactions. Prerequisite: OCEAN 520 or permission of instructor. Offered: A.
Instructor Course Description: James W Murray

OCEAN 522 Marine Organic Geochemistry (3)
Sources, reactions, and fates of organic molecules in the marine environment along with the stable isotope geochemistry of marine organic substances. Prerequisite: CHEM 237 and CHEM 239 or permission of instructor.
Instructor Course Description: Anitra E. Ingalls

OCEAN 523 Geochemical Cycles (4)
Descriptive, quantitative aspects of earth as biogeochemical system. Study of equilibria, transport processes, chemical kinetics, biological processes; their application to carbon, sulfur, nitrogen, phosphorus, other elemental cycles. Stability of biogeochemical systems; nature of human perturbations of their dynamics. Prerequisite: permission of instructor. Offered: jointly with CHEM 523 and ATM S 508.

OCEAN 524 Environmental Chemical Modeling (3) Benjamin, Murray
Physical/chemical principles controlling the fate and distribution of environmental pollutants, and use of models to apply those principles. Includes modeling of physical transport in conjunction with chemical equilibrium and reaction kinetics. Applications include acid mine drainage, acid deposition, and groundwater and lake water contamination. Offered: jointly with CEE 550.

OCEAN 529 Seminar on Chemical Oceanography (*, max. 9)
Lectures, discussions, and readings on selected problems of current interest. Prerequisite: permission of instructor. Offered: AWSp.

OCEAN 530 Biological Oceanography: Bacteria and Protozoa (3)
Bacteria in the marine environment; fate of organic carbon in the ocean and the interrelationship of the carbon cycle with other biogeochemical cycles. Prerequisite: permission of instructor. Offered: W.

OCEAN 531 Biological Oceanography: Phytoplankton (3)
Phytoplankton in the marine environment: ecology, primary productivity, and physiology. Phytoplankton growth and photosynthetic patterns; spatial and temporal distributions of phytoplankton; methods for determining distributions and rates of production and growth. Prerequisite: permission of instructor. Offered: W.

OCEAN 532 Biological Oceanography: Zooplankton (3)
Distribution and abundance of pelagic animals in space and time; analysis of their interactions. Small-scale distributions and behavior, population dynamics and energetics, trophic structure and dynamics, pelagic community structure, models of populations and food chains, secondary production and biogeography. Prerequisite: permission of instructor. Offered: Sp.
Instructor Course Description: Julie E Keister

OCEAN 533 Biological Oceanography: Benthos (3)
Analysis of marine benthic communities; new research questions and method; ecologically important physics of benthic boundary layer; theories, mechanics, and observations of deposit feeding; succession as consequence of physical processes and biological interactions. Environments include deep-sea, continental shelves, estuaries, and intertidal, focusing on soft substrata. Prerequisite: permission of instructor. Offered: Sp.

OCEAN 534 Methods and Measurements in Biological Oceanography (2)
Methods for bacteria, phytoplankton, and zooplankton population assessment. Rate measurements of phytoplankton, zooplankton, and bacterial production. Benthos measurements, including deep-sea environments. Prerequisite: permission of instructor.

OCEAN 535 Biological Oceanography: An Overview (3)
Principles and practice of biological oceanography for students with strong background in physical sciences but little recent exposure to biology. Ecological principles at individual, population, and community levels; overview of discipline of biological oceanography; case studies of interdisciplinary problems shared with the physical sciences. Offered: W.

OCEAN 536 Seminar in Geostatistics (1-3, max. 3)
Lectures and discussions on selected problems in the applications of statistics in earth science.

OCEAN 539 Seminar in Biological Oceanography (*, max. 9)
Lectures, discussions, and work on selected problems of current interest. Prerequisite: permission of instructor. Offered: AWSp.
Instructor Course Description: Daniel Grunbaum

OCEAN 540 Marine Geological Processes (3) McDuff
Principles of thermodynamics, heat and mass transfer, fluid mechanics, continuum mechanics, and time-series analysis applied to marine geological and geophysical data with special applications to thermal balance of the oceanic lithosphere. Offered: W.

OCEAN 541 Marine Sedimentary Processes (3)
Investigates fundamental process of marine sedimentation, including equations characterizing boundary-shear flows, initiation of grain motion, bedload and suspended-load transport, and sediment accumulation. Applies concepts to sediment dispersal in rivers, deltas, estuaries, beaches, continental shelves, slopes, and rises, with emphasis on the relationships between active processes and resulting deposits.

OCEAN 542 Sediment Dynamics and Boundary-Layer Physics (4)
Theoretical descriptions of sediment transport processes constrained by laboratory demonstrations. The physics of boundary layers, initiation of motion, suspended load, bedload, bedforms, and continua transport (turbidity currents, debris flows, and suspensions) and its application to the geological record. Offered: jointly with ESS 526; W.
Instructor Course Description: Jeffrey D Parsons

OCEAN 545 Physics of the Oceanic Lithosphere (3)
Basic principles of elasticity, fluid flow, and heat transport with specific applications to the formation and evolution of the oceanic lithosphere. Includes deformation of the earth, flow in porous media, heat transport, and marine seismological and potential field techniques. Prerequisite: OCEAN 540. Offered: jointly with ESS 568.

OCEAN 546 Continental-Margin Sedimentation (3)
Detailed evaluation of recent studies into processes forming strata on continental margins, including the diverse time scales ranging from sediment transport to sequence stratigraphy. Highlights the linkages with physical oceanographic processes, the fates of geochemical components, and the relationship to biological communities. Offered: jointly with ESS 527.

OCEAN 549 Seminar in Geological and Geophysical Oceanography (*, max. 9)
Lectures, discussions, and field and laboratory work on selected problems of current interest. Prerequisite: permission of instructor. Offered: AWSp.
Instructor Course Description: Russell E Mc Duff

OCEAN 550 Geochemistry and Geophysics of Melt Generation (3)
Mantle flow beneath mid-ocean ridges and hotspots, major element systematics, constraints from trace elements and isotopes on melting and mantle reservoirs, melt extraction, and crustal thickness and axial topography. Prerequisite: OCEAN 544 or permission of instructor. Offered: alternate years.

OCEAN 551 Marine Seismology (3)
Practical application of seismic techniques to the study of the ocean basins. Analysis of refraction data, multichannel reflection profiling, surface wave studies, and earthquake analysis. Prerequisite: GPHYS 502 or permission of instructor.

OCEAN 552 Seminar in Geophysics and Geological Data Analysis (1)
Practical geophysical data analysis, map projections, gridding multibeam bathymetry processing, gravity and magnetic anomalies, downward continuation, magnetic inversion, seismic refraction and reflection, and microearthquake locations. Prerequisite: permission of instructor.

OCEAN 558 Climate Modeling (3) Bitz, Thompson
Principles of Earth system modeling. Emphasis on atmosphere, ocean sea ice, and land-surface components. Climate forcing. Appropriate use of models. Topics of current interest including carbon cycle, atmosphere chemistry, and biogeochemistry. Prerequisite: either ATM S/OCEAN/ESS 587, ATM S 504 or ATM S 505. Offered: jointly with ATM S 559/ESS 559; Sp; alternate years.

OCEAN 559 Advanced Seminar on Mid-Ocean Ridge Processes (*, max. 9)
Lectures, discussions, and practical work on selected topics of current interest in mid-ocean ridge research. Prerequisite: permission of instructor.

OCEAN 560 Atmosphere/Ocean Interactions (3)
Observations and theory of phenomena of the coupled atmosphere-ocean system. El Nino/Southern Oscillation; decadal tropical variability; atmospheric teleconnections; midlatitude atmosphere-ocean variability. Overview of essential ocean and atmospheric dynamics, where appropriate. Credit/no credit only. Prerequisite: ATM S 509 or OCEAN 512. Offered: jointly with ATM S 560; alternate years; Sp.

OCEAN 569 Topics in Physical Oceanography (1-4, max. 9)
Lecture series on topics of major importance in physical oceanography. Offered: AWSp.
Instructor Course Description: Kevin L. Williams

OCEAN 570 Marine Microbial Interactions (1-3, max. 9)
Structure, function, and dynamics of natural mixed-species populations of marine bacteria and their interactions with higher organisms; mixed-species culture methods; synecological field methods; species assemblages in specialized environments; mutualisms; sites and patterns of genetic exchange. Prerequisite: OCEAN 530 or permission of instructor. Offered: alternate years.

OCEAN 571 Marine Primary Productivity (1-3, max. 9)
Patterns and mechanisms of marine phytoplankton primary production. Small-to-global-scale patterns of production; environmental regulation of production; absorption of electromagnetic radiation; fluorescence; carbon fixation; trophic interactions; remote sensing and other optical methods. Prerequisite: OCEAN 531 or permission of instructor. Offered: alternate years.

OCEAN 572 Zooplankton Ecology (1-3, max. 9)
Life history strategies, dynamics and production of populations, vertical migration, interspecific interactions and community structure, models of complex assemblages of zooplankton, sampling methods and analysis, spatial heterogeneity. Prerequisite: OCEAN 532 or permission of instructor. Offered: alternate years.

OCEAN 573 Benthic Biological Processes (1-3, max. 9)
Processes characteristic of soft-bottom benthic environments; areas and methods of rapid current progress; open research questions; deposit feeding; passive larval recruitment; physical, chemical, geological, and biological feedbacks in ecological succession; scaling of laboratory systems. Prerequisite: OCEAN 533 or permission of instructor. Offered: alternate years.

OCEAN 574 Principles and Applications of Molecular Methods (3)
Applications of molecular techniques to questions in ecology, evolution, and natural resource management, with emphasis on advanced genome-enabled technologies and the analysis and interpretation of genetic data. Includes weekly discussion of empirical papers, and preparation/peer-review of a proposal Prerequisite: permission of instructor. Offered: jointly with FISH 542; A.
Instructor Course Description: Lorenz Hauser

OCEAN 575 Molecular Techniques (4)
Laboratory on DNA methods. Experiments analyzing genetic variation at the intra- and interspecific level, including one experiment of student's own design. Techniques include DNA extraction and quantitation, PCR, DNA sequencing, RFLP analysis and cloning. Prerequisite: FISH 542 or OCEAN 574 or permission of instructor. Offered: jointly with FISH 543; W.

OCEAN 578 Advanced Topics in Biological Oceanography (*, max. 18)
Specialized research areas. Topic varies each year. Offered at Friday Harbor Laboratories. Prerequisite: permission of director of Friday Harbor Laboratories. Offered: S.

OCEAN 580 Aquatic Kinetics (3)
Reaction rates and mass transport in water. Theories of chemical kinetics; experimental results from: CO2 hydrolysis, Fe, Mn, and H2S oxidation, stable isotope fractionation, mineral dissolution; homogeneous, heterogeneous, microbial catalysis; reaction and transport at air-water, sediment-water, and O2/H2S interfaces. Prerequisite: permission of instructor.

OCEAN 581 Geochemical Modeling (3)
Background to modeling concepts frequently encountered in chemical oceanography: box models, advection-diffusion problems, sediment diagenesis equations, and boundary layer (air-water and sediment-water interface) models. Problems requiring application of the models to chemical distributions in the ocean. Prerequisite: permission of instructor.

OCEAN 582 River Basin Biogeochemistry (3)
The function of rivers and river basins in transporting materials to the oceans and their importance in biogeochemical cycles. Origin of water and water routing within drainage basins, sources and modification of dissolved and particulate materials in transport, ecological theory, and estuarine mixing zone transformations. Prerequisite: permission of instructor.

OCEAN 583 Isotope Biogeochemistry (3)
The use of stable isotopes to study biogeochemical cycles in the oceans and atmosphere; specifically carbon, nitrogen, and sulfur cycles. Isotopic effects during photosynthesis, respiration, organic matter degradation. CaCO3 dissolution, methanogenesis, nitrification/denitrification, and sulfate reduction. Prerequisite: permission of instructor.
Instructor Course Description: Paul D Quay

OCEAN 584 Ocean Tracers and Mixing (3)
The applications of tracers to studies of ocean circulation and ventilation. Processes within the ocean for which tracers have provided important information include gas exchange, mixed layer dynamics, thermocline ventilation, deep water formation and spreading, and mixing. Knowledge of partial differential equations suggested.

OCEAN 585 Paleoceanography (3)
History of environmental changes on earth over the past 100 million years as reconstructed from records in deep-sea sediments, ice sheets, and other ocean/terrestrial substrates. Examination of isotopic, geochemical, micropaleontological, and dating techniques. Role of the ocean in climate change. Prerequisite: permission of instructor.

OCEAN 586 Current Research in Climate Change (2, max. 20)
Weekly lectures focusing on a particular aspect of climate (topic to change each year) from invited speakers (both UW and outside), plus one or two keynote speakers, followed by class discussion. Offered: jointly with ATM S 586/ESS 586.
Instructor Course Description: James W Murray

OCEAN 587 Climate Dynamics (3) Hartman, Thompson
Examines Earth's climate system; distribution of temperature, precipitation, wind ice, salinity, and ocean currents; fundamental processes determining Earth's climate; energy and constituent transport mechanisms; climate sensitivity; natural climate variability on interannual to decadal time scales; global climate models; predicting future climate. Offered: jointly with ATM S 587/ESS 587. Offered: A

OCEAN 588 The Global Carbon Cycle and Climate (3) Emerson
Oceanic and terrestrial biogeochemical processes controlling atmospheric CO2 and other greenhouse gases. Records of past changes in the earth's carbon cycle from geological, oceanographic and terrestrial archives. Anthropogenic perturbations to cycles. Develop simple box models, discuss results of complex models. Offered: jointly with ATM S 588/ESS 588. Offered: W.

OCEAN 589 Paleoclimatology: Data, Modeling and Theory (3) Battisti, Emerson, Steig
Evidence for past changes in land and se surface temperature, in precipitation nd atmospheric dynamics, and in ocean circulation: both long and interannual timescales. Paleoclimate modeling and theory. Time series analysis and climate noise. Rapid climate change. Statistical reconstruction of interannual variability. Offered: jointly with ATM S 589/ESS 589. Offered: Sp.

OCEAN 590 Advanced Topics in Oceanography (9-18, max. 18)
Advanced topics examining specialized and interdisciplinary areas of oceanographic research. Offered at Friday Harbor Laboratories. Prerequisite: permission of Director of Friday Harbor Laboratories. Offered: S.

OCEAN 591 Marine Science in the Coastal Zone (3) Klinger
Presentation and analysis of the marine science of estuarine, coastal, and open ocean systems, including evaluation and interpretation of scientific information necessary for management. Lectures, discussions, and readings emphasize the relevance of natural processes to marine environmental management and decision-making. Offered: jointly with SMA 591; A.

OCEAN 592 Communicating Ocean Sciences (4) Robigou
Explores improving communicating scientific knowledge through instruction of inquiry-based science in school classrooms. Students practice communicating science, and receive mentoring on how to improve their presentations. Prerequisite: Marine sciences studies backgroung. Offered: Sp.

OCEAN 600 Independent Study or Research (*)
Offered: AWSpS.

OCEAN 700 Master's Thesis (*)
Offered: AWSpS.

OCEAN 800 Doctoral Dissertation (*)
Offered: AWSpS.